2011年3月17日木曜日

Ch.15 Tracing Evolutionary History

WHAT ARE THE DIFFERENCES BETWEEN NATURAL SELECTION AND EVOLUTION?
Evolution is a gradual process in which something changes into a different and usually more complex or better form whereas natural selection is the process in nature by which, according to Darwin's theory of evolution, only the organisms best adapted to their environment tend to survive and transmit their genetic characteristics in increasing numbers to succeeding generations while those less adapted tend to be eliminated.

HOW DOES EVOLUTION OCCUR?
Evolution is the change in frequency of the alleles in a gene pool over time. Evolution occurs with mutations and natural selections. WHat happens is that in a population of species, not all of them are the same. For example, there is a whole colony of fish. Some of the fish are faster than the others. They reproduce and pass on their genes which make their offspring fast as well. The other fish that aren't as fast, die out because they are not fast enough. The fast fish soon become more common, making the species as a whole faster. This occurs in several stages, where the slowest fish are removed each time, and the faster ones reproduce more often. The second aspect is that these fish will often have differences in their genetic codes, called mutations. While bad mutations will normally result in fatality and not get passed on, some are useful and result in increased survival. When these mutations build up, the species we previously had may then become completely different from what is here now. The two groups cannot cross breed, and have become different species.

WHAT TYPE OF EVOLUTIONARY EVENT HAS OCCURRED?
Natural selection and a genetic bottleneck. The DDT resistance is a classic example of natural selection, only the most resistant organisms surviving the treatment and establishing a new, resistant population. The near disappearance of the mosquito population for several years with the new population showing reduced fertility is characteristic of a genetic bottleneck. Only a very small number survived and the genetic diversity in the population is greatly reduced. The survivors are forced to undergo massive inbreeding and reduced fertility is quite common.


SUMMARY:
A recipe for life is raw materials, suitable environment, and energy sources. The conditions on early Earth made the origin of life possible. The chemical and physical conditions made abiotic synthesis of monomers (1st stage), formation of polymers (2nd stage), packaging of polymers into protobionts (3rd stage), and self-replication (4th stage.) In 1953, Stanley Miller tested the Oparin-Haldane hypothesis. He set up an airtight apparatus with gases circulating past an electrical discharge, to stimulate conditions on the early Earth. An alternative hypothesis states that submerged volcanoes and deep-sea hydrothermal vents may have provided the chemical resources for the first life. Cells and protobionts share four characteristics: structural organization, simple reproduction, simple metabolism, and simple homeostasis. The eukaryotic cell probably originated as a community of prokaryotes, when small prokaryotes capable of aerobic respiration or photosynthesis began living in larger cells. Oldest fossils of eukaryotes are about 2 billion years old. Sliding plates are earthquake zones. Also colliding plates form mountains. Those are geologic processes occur at plate boundaries. The supercontinent pangaea altered habitats and triggered the greatest mass extinction in Earth's history. Extinction is the fate of all species and most lineages. The history of life reflects a steady background extinction rate with episodes of mass extinction. Types of extinctions are classified by how shallow/deep water the species lived in. For example, permian extinction, cretaceous extinction.
Human development is paedomorphic, retaining juvenile traits into adulthood. In the evolution of an eye or any other complex structure, behavior, or biochemical pathway, each step must bring a selective advantage to the organism possessing it and must increase the organism's fitness. Other novel structures result from exaptation, the gradual adaptation of existing structures to new functions Natural selection doesn't anticipate the novel use: each intermediate stage must be adaptive and functional. Evolutionary trends do not mean that evolution is goal directed. Natural selection results from the interactions between organisms and their environment. If the environment changes, apparent evolutionary trends may cease or reverse. Systematics classifies organisms and determines their evolutionary relationship. Taxonomists assign each species a binomial consisting of a genus and species name. A phylogenetic tree is a hypothesis of evolutionary relationships within a group. Cladistics uses shared derived characters to group organisms into clades, including an ancestral species and all its descendents. Shared ancestral characters were present in ancestral groups. Molecular systematics compares nucleic acids or other molecules to infer related ness of taxa.


KEY TERMS:
- Macroevolution: major changes over evolutionary time
- Radiometric dating: measurement of the decay of radioactive isotopes
- Geologic record: defined by major transitions in life on Earth
- Continental drift: the slow, continuous movement of Earth's crustal plates on the hot mantle
- Adaptive radiation: a group of organisms forms new species, whose adaptations allow them to fill new habitats or roles in the communities
- Evo-devo: a field that combines evolutionary and developmental biology
- Homeotic genes: master control genes that determine basic features, such as where pairs of wings or legs develop on a fruit fly
- Species selection: the unequal speciation or unequal survival of species on a branching evolutionary tree
- Phylogeny: the evolutionary history of a species or group of species
- Convergent evolution: where analogous similarities result from in similar environments









This is a famous picture of evolution. It started out with the ape on the left, and ends with human being now on the right. It didn't happen just at once, but it took couple steps in between to become a human being from an ape. Also it took extremely long time. Like I said in the beginning, some apes who have different and better genes than others keep being alive, and other apes died out because of some reasons. And whoever that is still alive keep raising their children which are more likely to have "better" genes. This is how evolution occurs. This shows a microevlution.

http://www.youtube.com/watch?v=fVPadCMKJ_8


5 FACTS:
1) Homologous genes have been found in organisms separated by huge evolutionary distances
2) Molecular clocks help track evolutionary time. It can be calibrated in real time by graphing the number of nucleotide differences against the dates of evolutionary branch points known from the fossil record.
3) Life is divided into three domains: the prokaryotic domains Bacteria and Archaea and the eukaryote domain Eukarya.
4) There have been two major episodes of horizontal gene transfer, with transfer of genes between genomes by plasmid exchange, viral infection, and fusion of organisms: gene transfer between a mitochondrial ancestor and the ancestor of eukaryotes, and gene transfer between a chloroplast ancestor and the ancestor of green plants.
5) Shared characters are used to construct phylogenetic trees.

2011年3月2日水曜日

Ch.14 The Origin of Species

WHY CAN'T MULES MATE?
The basic reason why mules cannot reproduce is that a mule does not have a even number of chromosomes. They have 63 chromosomes because a horse and a donkey have different numbers of chromosomes. During mitotic cell division, each of the chromosomes copies itself and then distributes these two copies to the two daughter cells. In contrast, when the mule is producing sperm or egg cells during meiosis, each pair of chromosomes need to pair up with each other. Since the mule doesn't have an even number of homologous pairs, meiosis is disrupted and viable sperm and egg are not formed.

WHAT ARE PREZYGOTIC REPRODUCTIVE BARRIERS?
There are two general categories of reproductive isolating mechanisms: prezygotic, or those that take effect before fertilization, and postzygotic, hybrids between members of different populations through ecological, temporal, ethological, mechanical, and gametic isolation.

WHAT IS THE PHYLOGENETIC SPECIES CONCEPT?
The phylogenetic species concept defines a species as a set of organisms representing a specific evolutionary lineage. The concept of a species as an irreducible group whose members are descended from a common ancestor and who all possess a combination of certain traits. It is less restrictive than the biological species concept. Also it permits successive species to be defined even if they have evolved in an unbroken line of descent, with continuity of sexual fertility. However because slight differences can be found among virtually any group of organisms, the concept tends to encourage extreme division of species into ever-smaller groups.


SUMMARY:
Linnaeus used physical characteristics to distinguish species. His binomial system is the basis of taxonomy, the naming and classification of life's forms. The biological species concept defines a species as a group of populations whose members can interbreed and produce fertile offspring with each other but not with members of other species. Most organisms are classified based on observable phenotypic traits; the morphological species by its ecological niche. According to the phylogenetic species concept, a species is the smallest group that shares a common ancestor and forms one branch on the tree of life.
There are two types of reproductive barriers. Prezygotic barriers, which includes temporal isolation, habitat isolation, behavioral isolation, mechanical isolation, and gametic isolation. Also postzygotic barriers, which includes reduced hybrid viability, reduced hybrid fertility, and hybrid breakdown.
Geographically separated from other populations, a small population may become genetically unique as its gene pool is changed by natural selection, mutation, or genetic drift in allopatric speciation. A laboratory study has documented the beginning of reproductive isolation as fruit fly populations adapted to a new food source. Researchers have identified the specific genes involved in some cases of speciation, which is the evolution of reproductive barriers. Hybrid zones are regions in which members of different species overlap and produce at least some hybrid offspring. Over time, reinforcement may strengthen prezygotic barriers to reproduction, or fusion may reverse the speciation process as reproductive barriers weaken and extensive gene flow occurs. In stable hybrid zones, a limited number of hybrid offspring continue to be produced. Adaptive radiation can occur when populations are provided with expanded opportunities following mass extinctions, the colonization of a diverse new environment, or the evolution of new structures. The punctuated equilibrium model draws on the fossil record, where many species change most as they arise from an ancestral species and then change relatively little for the rest of their existence. But some species have evolved by the gradual accumulation of changes. The time interval between speciation events varies considerable, from a few thousand years to tens of millions of years.



white-horse_wallpapers_4836_1024x768.jpg     +    donkey.png   =
Mule.jpg


A Mule is a hybrid of a horse and a donkey. However mules cannot mate. Therefore a mule is an animal that never reproduce. A mule is not considered as a specie, because species have to be able to inbreed. In order to make a mule, a horse and a donkey needs to mate.

http://www.youtube.com/watch?v=zl1a1n6XR9g

KEY TERMS:
- Species: as a group of organisms whose members can breed and produce fertile offspring, but who do not produce fertile offspring with members of other groups
- Taxonomy: the branch of biology that names and classifies species and groups them into broader categories
- biological species concept: species as a group of populations whose members have the potential to interbreed in nature and produce fertile offspring
- Morphological species concept: classification is based mainly on phenotype
- Ecological species concept: identification of species in terms of their ecological niches, focusing on unique adaptations to particular roles in a biological community
- Phylogenetic species concept: a species as the smallest group of individuals that shares a common ancestor and that forms one branch on the tree of life.
- Reproductive barrier: a biological feature of the organism itself
- Allopatric speciation: populations separated by a geographic barrier
- Polyploid: their cells have more than two complete sets of chromosomes
- Sympatric speciation: a new species arises within the same geographic area as a parent species

5 FACTS:
1) Reproductive barriers keep species separate
2) In allopatric speciation, geographic isolation leads to speciation
3) In sympatric speciation, speciation takes place without geographic isolation
4) Reproductive barriers may evolve as populations diverge
5) Hybrid zones provide opportunities to study reproductive isolation